While on a camping trip with Uncle Louie, two young mice named Angelina and Henry become lost in the woods until the "fit and fearless" Angelina saves the day.
Angelina finds out just how important a lucky penny can be--and whether it can take the place of practice--as she prepares to audition for a role at the Theater Royale.
Angelina Ballerina is upset when her young cousin Henry is chosen to perform with the famous Madame Zizi, but she is soon rewarded for her efforts to help him learn the steps of the dance.
Angelina is eager to have her friend Anya Moussorsky from Dacovia stay with her for the summer and attend ballet classes, but she lets jealousy get the best of her when it seems Anya is getting too much attention.
Angelina, headed to the Mouseland Dance Festival aboard her grandparents' canal boat, is so obsessed with practicing her dance moves, and so neglectful of her duties, that the trip turns into a near disaster.
When Angelina does not get the star role in the Christmas dance recital, she becomes upset and quits the show, but when one of the performers is injured she suddenly sees the importance of doing the role given her to her best ability.