Woolly mammoth siblings Oscar and Arabella enjoy being at Mammoth Academy, but Oscar is accused of stealing oranges and when he follows some mysterious tracks to find the real thief, he discovers humans living nearby.
Ayla and Jondalar meet the Mautoi, the Mammoth Hunters, people like Ayla. Ayla finds herself torn between the wildly jealous Jondalar and the master carver Ranec.
Sam Monroe, the reluctant commander of a tough-minded warrior people living in what was once northern Mexico, must fight the Khan forces before they conquer his troops and his kingdom.
Manny, Sid, and Diego, along with new friends Ellie, Crash, and Eddie must find a way to escape the oncoming flood when the ice around them begins to melt.