After entering a great stone structure and escaping deadly Ugha warriors, Indiana Jones realizes they're not in the true Lost City. He and his company operate an ancient mechanism to reveal Akator and the mysteries the Ugha have been protecting.
Spalko, Dovchenko, and Mac have captured Indy's group and are heading through the Amazon, but Indy, Mutt, and Marion escape to the jungle and are soon joined by Ox, who is desperate to return the crystal skull to Akator.
Indiana Jones and Omphale attempt to flee from the Cult of Hecate in a stole German war plane and try to stay one step ahead of those seeking the fabled Golden Fleece.