Angelina is thrilled when she receives the starring role in the new ballet until she learns she needs to fly across the stage dangling from a single wire.
When Cousin Henry gets all the attention in the grown-up musical in which they have roles, Angelina becomes jealous, but she demonstrates her fondness for him when a crisis occurs on opening night.
Angelina insists she wants Anya, a mouseling visiting from Dacovia for the summer, to stay at her house and share her room, but Angelina's feelings change when Anya begins getting too much attention.
Angelina and the other mouselings at Miss Lilly's Ballet School perform Cinderella on tour, battling homesickness and bad luck to ensure the show will go on.